Once upon a time in the far away kingdom of Dor lived a brave and virtuous mouse with comically oversized ears who dreamt of becoming a knight. Banished from his home for having such lofty ambitions, Despereaux sets off on an amazing adventure with his good-hearted rat friend Roscuro. The two go on a very noble quest to rescue an endangered princess and save an entire kingdom from darkness.

A young girl walks through a secret door that she has found in her new home and discovers an alternate version of her life. On the surface, this parallel reality is eerily similar to her real life, but much better. When her adventure turns dangerous, and her counterfeit parents, including the Other Mother, try to keep her forever, Coraline must count on her resourcefulness, determination, and bravery to get back home - and save her family.